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Edinger-westphal Nucleus Information

The Edinger-Westphal nucleus (also known as the accessory oculomotor nucleus ) is the accessory parasympathetic cranial nerve nucleus of the oculomotor nerve (cranial nerve III), supplying the constricting muscles of the iris. Alternatively, the Edinger-Westphal nucleus is a term often used to refer to the adjacent population of non-preganglionic neurons that do not project to the ciliary ganglion, but rather project to the spinal cord, dorsal raphe nucleus, and lateral septal nuclei. Unlike the classical, preganglionic Edinger-Westphal neurons that contain choline acetyltransferase, neurons of the non-preganglionic Edinger-Westphal nucleus have been shown to contain various stress- and feeding-related neuropeptides, such as Urocortin and cocaine- and amphetamine-regulated transcript.[1] Recently a proposal to rename this group of non-preganglionic, neuropeptide-containing neurons to perioculomotor subgriseal neuronal stream, abbreviated pIIISG, has been suggested.[2]

Contents

Location

The paired nuclei are posterior to the main motor nucleus (oculomotor nucleus) and anterolateral to the cerebral aqueduct in the rostral midbrain at the level of the superior colliculus.

It is the most rostral of the parasympathetic nuclei in the brain stem.

Function

The Edinger-Westphal nucleus supplies preganglionic parasympathetic fibers to the eye, constricting the pupil, accommodating the lens, and convergence of the eyes.[3]

It has also been implicated in the mirroring of pupil size in sad facial expressions. When seeing a sad face, participants' pupils dilated or constricted to mirror the face they saw, which predicted both how sad they perceived the face to be, as well as activity within this region.[4][5]

Eponym

The nucleus is named for both Ludwig Edinger, from Frankfurt, who demonstrated it in the fetus in 1885, and for Karl Friedrich Otto Westphal, from Berlin, who demonstrated it in the adult in 1887.[6]

Additional images

The cranial nerve nuclei schematically represented; dorsal view. Motor nuclei in red; sensory in blue.

Nuclei of origin of cranial motor nerves schematically represented; lateral view.

Primary terminal nuclei of the afferent (sensory) cranial nerves schematically represented; lateral view.

References

  1. ^ Kozicz, T. (2003). "Neurons colocalizing urocortin and cocaine and amphetamine-regulated transcript immunoreactivities are induced by acute lipopolysaccharide stress in the Edinger-Westphal nucleus in the rat". Neuroscience 116 (2): 315–20. doi:10.1016/S0306-4522(02)00772-8. PMID 12559087.
  2. ^ May PJ, Reiner AJ, Ryabinin AE (March 2008). "Comparison of the distributions of urocortin-containing and cholinergic neurons in the perioculomotor midbrain of the cat and macaque". J. Comp. Neurol. 507 (3): 1300–16. doi:10.1002/cne.21514. PMID 18186029.
  3. ^ "eye, human" Encyclopædia Britannica. Encyclopaedia Britannica Ultimate Reference Suite. Chicago: Encyclopædia Britannica, 2010.
  4. ^ Harrison, NA; Wilson, CE; Critchley, HD (2007). "Processing of observed pupil size modulates perception of sadness and predicts empathy.". Emotion (Washington, D.C.) 7 (4): 724–9. doi:10.1037/1528-3542.7.4.724. PMID 18039039.
  5. ^ Harrison, NA; Singer, T; Rotshtein, P; Dolan, RJ; Critchley, HD (2006). "Pupillary contagion: central mechanisms engaged in sadness processing.". Social cognitive and affective neuroscience 1 (1): 5–17. doi:10.1093/scan/nsl006. PMID 17186063. PMC 1716019. http://www.pubmedcentral.nih.gov/picrender.fcgi?artid=1716019&blobtype=pdf.
  6. ^ synd/893 at Who Named It?

External links

· · Human brain: mesencephalon (midbrain) (TA A14.1.06, GA 9.800)
Tectum (Dorsal)
Surface Corpora quadrigemina: Inferior colliculi (Brachium of inferior colliculus), Superior colliculi (Brachium of superior colliculus)
Grey matter Pretectal area
White: Sensory/ascending Spinotectal tract · Central tegmental tract
White: Motor/descending Tectospinal tract
Peduncle (Ventral)
Tegmentum
White: Sensory/ascending lemnisci (Medial, Lateral) · Ascending MLF (Vestibulo-oculomotor fibers) · Spinothalamic tract · Anterior trigeminothalamic tract · Dentatothalamic tract
White: Motor/descending Rubrospinal tract · Rubro-olivary tract · Descending MLF
Grey: cranial nuclei GSA (V: Mesencephalic) - GSE (III: Oculomotor, IV: Trochlear) - GVE (III: Edinger-Westphal)
Grey: other

Periaqueductal gray/Raphe nuclei (Dorsal raphe nucleus)

Ventral tegmental areaPedunculopontine nucleusRed nucleus

riMLF
Ventricular system Cerebral aqueduct
Base
White: Motor/descending Cerebral crus: Corticospinal tract · Corticobulbar tract · Corticopontine tract/Frontopontine fibers/Temporopontine fibers
Grey: Substantia nigra Pars compacta · Pars reticulata
Surface Superior cerebellar peduncle (Decussation of superior cerebellar peduncles) · Interpeduncular fossa

: CNS

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· · Nerves of head and neck: the cranial nerves and nuclei (TA A14.2.01, GA 9.855)
olfactory (AON->I) olfactory bulb · olfactory tract
optic (LGN->II) optic chiasm · optic tract
oculomotor (ON, EWN->III) superior branch (parasympathetic root of ciliary ganglion/ciliary ganglion) · inferior branch
trochlear (TN->IV) no significant branches
trigeminal (PSN, TSN, MN, TMN->V) trigeminal ganglionophthalmicmaxillarymandibular
abducens (AN->VI) no significant branches
facial (FMN, SN, SSN->VII)
near origin nervus intermediusgeniculate
inside facial canal greater petrosal (pterygopalatine ganglion) · nerve to the stapedius · chorda tympani (lingual nerve, submandibular ganglion)
at stylomastoid foramen posterior auricular · suprahyoid (digastric, stylohyoid) · parotid plexus (temporal, zygomatic, buccal, mandibular, cervical)
vestibulocochlear (VN, CN->VIII) cochlear (striae medullares, lateral lemniscus) • vestibular (Scarpa's ganglion)
glossopharyngeal (NA, ISN, SN->IX)
before jugular fossa ganglia (superior, inferior)
after jugular fossa tympanic (tympanic plexus, lesser petrosal, otic ganglion) • stylopharyngeal branchpharyngeal branchestonsillar brancheslingual branchescarotid sinus
vagus (NA, DNVN, SN->X)
before jugular fossa ganglia (superior, inferior)
after jugular fossa meningeal branch · auricular branch
neck pharyngeal branch (pharyngeal plexus) · superior laryngeal (external, internal) · recurrent laryngeal (inferior) · superior cervical cardiac
thorax inferior cardiac · pulmonary · vagal trunks (anterior, posterior)
abdomen celiac · renal · hepatic · anterior gastric · posterior gastric
accessory (NA, SAN->XI) cranial · spinal
hypoglossal (HN->XII) lingual branches

: PNS

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· · Sensory system: Visual system and eye movement pathways
Visual perception 1° (Bipolar cell of Retina) → 2° (Ganglionic cell) → 3° (Optic nerveOptic chiasmOptic tractLGN of Thalamus) → 4° (Optic radiationCuneus and Lingual gyrus of Visual cortexBlobsGlobs)
Muscles of orbit
Tracking Smooth pursuit: Parietal lobe · Occipital lobe Saccade: Frontal eye fields Physiologic nystagmusFixation reflexPPRF
Horizontal gaze PPRFAbducens nucleusMLFOculomotor nucleusMedial rectus muscle
Vertical gaze Rostral interstitial nucleusOculomotor nucleus, Trochlear nucleusMuscles of orbit
Vestibulo-ocular reflex Semicircular canalVestibulocochlear nerveVestibular nucleiAbducens nucleusMLF (Vestibulo-oculomotor fibers) → Oculomotor nucleusMedial rectus muscle
Pupillary reflex
Pupillary dilation 1° (Posterior hypothalamusCiliospinal center) → 2° (Superior cervical ganglion) → 3° (Sympathetic root of ciliary ganglionNasociliary nerveLong ciliary nervesIris dilator muscle)
Pupillary light reflex (constriction) 1° (RetinaOptic nerveOptic chiasmOptic tractPretectal nucleus) → 2° (Edinger-Westphal nucleus) → 3° (Oculomotor nerveParasympathetic root of ciliary ganglionCiliary ganglion) → (4° Short ciliary nerves → Iris sphincter muscle)
Accommodation vergence 1° (RetinaOptic nerveOptic chiasmOptic tractVisual cortexBrodmann area 19Pretectal area) → 2° (Edinger-Westphal nucleus) → 3° (Short ciliary nervesCiliary ganglionCiliary muscle)
Circadian rhythm RetinaHypothalamus (Suprachiasmatic nucleus)

: EYE

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Categories: Neuroanatomy

 

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